Saturday, June 16, 2018

On Rediscovering "Extinct" Species: The Lost and Found Project ---Guest Post---

This is a guest post by Dr. Diogo VerĂ­ssimo at the University of Oxford, describing a project recently supported in part by The Alongside Wildlife Foundation.

It’s easy to feel a bit down if you follow environmental news from around the globe. Every day there seems to be a new tragedy, a more recent disaster, a more pressing crisis. This is not without reason. The natural world is under threat and in need of more attention, but perhaps there should also be room for some more inspirational stories? 

Photo by E.J. Keller Baker (Public Domain)
As a child, I was consumed by the story of the Thylacine or Tasmanian tiger, a predator marsupial found only in the island of Tasmania, in Australia. Although this is no doubt a spectacular animal, being the largest known carnivorous marsupial of modern times. It is different to any other existing animal on earth today, representing a unique branch of our planet’s tree of life. But what mesmerized me about this species was the uncertainty around its extinction. While the mainstream narrative is that the last Tasmanian tiger died in captivity in 1936, there have been dozens of sightings of this species, leading to several expeditions to attempt to rediscover it. What if this amazing creature was still out there? What if this unique animal was after all not lost forever? 

I started reading up on these cases, where a species thought extinct was eventually rediscovered. It seemed that the Tasmanian tiger was not alone, and while it seems increasingly unlikely that it will ever be rediscovered, other species have. Many of these rediscoveries are truly inspirational stories, not only of how some amazing biodiversity “came back from the dead”, but also of the incredible commitment, selflessness and courage of the scientists, park rangers and nature enthusiasts involved. Here was my antidote to the doom and gloom that dominates so much of the environmental news. The Lost & Found project was born.

On Earth Day 2017, at the Earth Optimism Summit in Washington DC, the Lost & Found project was launched, with the goal of telling the most surprising, exciting and serendipitous stories of species rediscoveries. We currently cover 20, soon to be 25 species, from craneflies to turtles and newts to flying squirrels, from five continents. We present our stories in both written format and comics to allow for different audiences to enjoy these rediscoveries.

Thanks to a grant from The Alongside Wild Foundation, we were recently able to make important changes to our website, including the addition of five more species and the possibility to translating the content to other languages. So what’s next? We are currently working on five new stories, this time featuring plants, and are hoping to raise funds to have video animations of each of our stories, opening up our content to even more people.

So if you haven’t heard the story of how David Wingate moved his family to a desert island to save the Bermuda Petrel, how the rediscovery of the Bulmer’s fruit bat almost did not happen due to the appetite of a dog, or how Peter Zahler had to resort to a local collector of squirrel poo to rediscover the Woolly flying squirrel, then I challenge you to pay us a visit and get to know some of these incredible tales.

Sunday, June 3, 2018

Announcing the Wildlife Grants Awarded by The Alongside Wildlife Foundation

   Hi all, in 2017 I signed up with Patreon to find more justification for all the time I was spending doing outreach and science communication and also because I lacked job security and I was testing the waters to see what paths forward I could create. Once the donations starting rolling in I created a new non-profit charitable foundation, The Alongside Wildlife Foundation, to formalize this work and so that donations would be tax-deductible. In November I gave notice and quit my job at Auburn University. My plan was to develop the foundation (and donate plasma, rent out rooms, do freelance writing, etc) as the next stage of my life. But, shortly thereafter I was offered an exciting new position at the Georgia Sea Turtle Center (I started in January).

   Importantly, because I finally have job security and life stability I was able to make some big changes with the foundation funds I originally planned to keep for my salary. Since January, all donations have been used to grow The Alongside Wildlife Foundation and help other scientists and science communicators accomplish the mission we all share: science-based solutions for living alongside wildlife.

   One of our major initiatives was a small grants program and thanks to our growing army of small donors we have already awarded about $4,000 this year to wildlife projects everywhere from Arkansas to Nigeria. I wanted to highlight the projects that have received funding so far here (and yes, perhaps entice you to become a recurring donor so we can start doing even more).

Ikponke Nkanta (Tropical Research and Conservation Centre): Primate inventory and conservation in Ikea River Basin, Southern Nigeria.
The goals of this project include ensuring 1) future survival of the threatened/important primate species occurring within Ikpa river basin in southern Nigeria through participatory habitat restoration, protection and management. 2) a reduction in forest land encroachment and over-dependence on forest resources as the major source of livelihood (by facilitating adoption of livelihoods alternatives (bee farming, snail farming, fish farming) among hunters and timber harvesters. 3) community adoption of sustainable timber harvesting and better farming practices that are habitat friendly (agro forestry, organic farming) as these do not support further forest destruction, and finally, 4) establishing buffer zones with indigenous trees around the present shrinking territories to stabilize and ‘bridge-up’ fragmented patches.

Lauren Hennelly (UC-Davis): Assessing the status and distribution of wolves in Pakistan using genetics.
The overall goal of this project is to provide important baseline information and resolve what types of wolves are found in Pakistan to inform conservation decisions with several components, including 1) conducting the first genetic survey of wolves across major regions of Pakistan, 2) providing estimates of genetic diversity and potential gene flow to adjacent wolf populations, and 3) provide a clearer picture of the distribution of the Central Indian and Himalayan wolf lineages and the extent of their range. Funds from The Alongside Wildlife Foundation are going towards the purchase of camera traps to gain more information on the local wolf populations and to document the phenotypic characteristics of the wolves to accompany the genetic data. Lastly, there are very few photographs of wild wolves from many regions of Pakistan – camera trap images can become educational resources for the local communities.

Karl and Diane Roeder (University of Oklahoma and Cameron University): The ants of Oklahoma project.
The core objective of the Ants of Oklahoma Project is to  ighlight the diversity of life that routinely goes unnoticed. To accomplish this goal, this program (1) combines historical records found in museum collections, published manuscripts, and online digital repositories with (2) a citizen science outreach component that involves collecting specimens in backyards using standardized methods. Cumulatively, this project will collate information on ant distributions across Oklahoma, create a database of these distributional records, and publish such information for all to view. Moreover, an outreach component will primarily focus on involving undergraduate students in field observations and identification of ants both throughout the year and during annual BioBlitz events. This multifaceted approach will not only increase awareness of our particular taxon of interest, ants (Hymenoptera: Formicidae), but will also encourage citizens and scientists alike to go outside, enjoy nature, and recognize how wonderfully diverse it truly is.

Diogo VerĂ­ssimo (University of Oxford): Lost and Found.

The storytelling project “Lost and Found” works to bring to life the inspirational stories of those species that were thought extinct for at least a human generation but where subsequently rediscovered. The goal of this project is to change the conversations around conservation, away from doom and gloom, and towards a more positive and balanced message about nature. The project will produce a total of 30 stories of serendipitous species rediscoveries across five continents and all animal and plant groups, both in narrative and comics format. This content is developed by professional science and creative writers and a comics artist and is freely available online. 

Wade Boys (University of Arkansas): Surveys, modeling, and prioritization for rare, endemic dragonflies across the Ozark-Ouachita region.

The goal of this project is to conduct rigorous field surveys for four dragonfly species of concern in the Ozark/Ouachita regions. All four of these species are rare and under-surveyed, with one of them being recently described. Their distributions are presumably limited by local habitat and climate characteristics, however, there’s been a lack of habitat-directed surveys. This study will address the sparse records of occurrence and unknown habitat requirements of these insects by combining distribution modeling with field surveys resulting in increased knowledge of their natural history.

Carla X. Neri Barrios (Soluciones Ambientales Itzeni, A.C., Mexico): A children’s book as a conservation education tool for awareness of the natural history of the lesser long-nosed bat (Leptonycteris yerbabuenae).
The goal of this project is to engage children of ages between 7 to 11 in the natural history of the lesser long-nosed bat (Leptonycteris yerbabuenae) through the production and distribution of a children's book with the aim of increasing awareness and empathy towards this and other bat species.

Zackary Graham (Arkansas State University): Field observations on the aggressive behavior of Ringed Crayfish in Colorado.

Despite the extensive research on crayfish physiology and ecology, their natural behavior is extremely understudied. It is often assumed that the behavior and aggression of all crayfish species are identical. But, it is commonly found that the species that have most successfully invaded other populations are more aggressive than less successful invaders. The goal of this project is to conduct behavioral observations of the Ringed Crayfish (Orcconectes neglectus), which is native to Colorado, Arkansas, and Missouri but has been introduced to several locations throughout North America. Currently, the aggressive nature of these crayfish is unknown, and their agonistic behavior has yet to be characterized. These observations will ultimately be used to understand how and why some crayfish are more successful invaders than others. Further understanding of crayfish natural history and specifically aggressive behavior are key to make informed conservation efforts.

Adam Mitchell (University of Delaware): Determining impacts of plant invasion on native arthropod diversity in the Mid-Atlantic.
The goal of this project is to quantify shifts in community structure for arthropods that are impacted by invasive plants in the Mid-Atlantic region. Specifically, it will 1) determine changes in the richness and abundance of arthropod functional groups (herbivores, detritivores, pollinators, and predators) following plant invasion and 2) investigate patterns of change in arthropod community structure in multiple invasive plants to identify which invasive plant (or plants) present the greatest threat to native arthropod diversity in the landscape. 

Erin Spencer (UNC-Chapel Hill): One fish, true fish: combatting seafood mislabeling in the United States.
Seafood fraud often allows less-desired or illegally-caught species to be marketed as a species popular with consumers.  Although there has been media coverage that seafood mislabeling exists, there are little to no resources to teach consumers how to avoid seafood fraud. The goal of this project is to create a website that contains information and tangible tools consumers can use to make smart, sustainable seafood choices that helps promote a culture that values accurate marketing of seafood. The website will include: 1) fact sheets derived from current mislabeling research, including which vendors types (like grocery stores, sushi restaurants, etc.) are most likely to mislabel, 2) web and PDF photo guides for how to identify common seafood replacements, and 3) blog posts and features of local fishermen and vendors trying to support honest, sustainable seafood.

Want to help us do even more to create and promote science-based solutions to living alongside wildlife in perpetuity? Please consider joining our growing network of recurring small donors here.

Wednesday, May 23, 2018

Is This a Wolf or Coyote? ---Guest Post---


Could you please identify as coyote or wolf.   

Thank you very much for your help!  

Mark S.

The identification between wolves and coyotes is often confused. The first thing to consider is their range. Historically, coyotes were restricted to the prairies that are west of the Mississippi River and into Central Mexico. However, due to logging, clear cutting for agriculture, the extirpation of the gray wolf and the rise of white-tailed deer populations in the 1800s, coyotes have expanded their range significantly. By the 1930s, they had dispersed around the Great Lakes and up into the northeastern states. Now they inhabit deserts, grasslands, mountains, forests, and rural and urban environments throughout the United States, Mexico, Central America, Canada and most of Alaska.

Map of Coyote range over time from Hody and Kays (2018)

Wolves currently range throughout Alaska, the Northern Rockies, Pacific Northwest, the Great Lakes region, and Canada. The Mexican gray wolf is a subspecies that inhabits the Apache National Forest in southeastern Arizona in a very small population of roughly 113 individuals.


Although at first glance wolves and coyotes can look similar, there are some characteristics that set these two distinct canids apart:

Photo courtesy of Macmanes
Coyotes weigh between 30-50 lbs, stand between 16-24 in. at the shoulder and are about 4 ft. long. Wolves weigh between 55-130 lbs, stand at 26-34 in. at the shoulder and are between 4.5-6 ft. long. Coat colors of coyotes vary from grayish tawny in the North, to a lighter tawny in its southern range. The underside and throat of both canids usually have buff or white coloring. The forelegs, sides of head and muzzle are a rusty shade on coyotes. Coyotes have a dorsal stripe formed by black-tipped guard hairs, as well as a black-tipped tail. The color variations of wolves range from white to black, and every shade of tawny in between with black-tipped guard hairs. Both coyotes and wolves have a black “diamond” marking on their tail up near the base that is where a scent gland is located. Coyotes are identified by their large, pointed ears, and narrow muzzle in comparison to wolves. Wolves have rounded ears and a broad muzzle. Both have long legs designed for endurance.

Photo Courtesy Tracy Brooks.
Northeastern coyotes are described as having more of a German Shepard-like appearance. Coyotes in this region are larger than those found anywhere else. This is attributed to the confirmation that northeastern coyotes share mitochondrial DNA (mtDNA) of eastern wolves. Many in the population also share mtDNA of domestic dog.

The animal in the photos sent in by Mark S, is of a coyote mother and her pups that appear to be about a month old. Breeding takes place between January and early March, and gestation last about two months. Pups are born between April and May into litters that average to around 4-5 pups. Den sites vary from rock ledges, hollow logs, and brush-covered slopes to depressions above the ground. Wolves have the same breeding period, gestation length, average litter size, and have similar den sites.

About the Author

Katey Duffey resides in Canton, Ohio, and has always had a passion for carnivore conservation. She has a B.A in Zoo and Wildlife Biology from Malone University, and a M.A in Zoology from Miami University. Katey has worked with a variety of species of reptiles and small mammals, and worked with captive and wild wolves. Currently, she is an independent researcher who has studied snow leopards in western Mongolia since 2014, with an interest in One Health that connects the wellbeing of a local community and their livestock with that of wildlife. She is also the Communications Director of The Tulsi Foundation, a UK-based organization that trains conservation staff and rangers in India trauma medicine where she uses her combat lifesaver experience from the Army and skills in wilderness medicine. 

Friday, May 4, 2018

Snake Identification Challenge of the Week - Identify These Snakes!

Greetings Dr., could you help ID the snake in photos attached?  I have combed many internet sites to no avail.  Found it entering my garage in central Oklahoma.  Since it does not match any Okla types, I'm thinking it might have been an escaped pet.
Thank you,
J. P.

Hi Doc,

I ran across this little guy in Greenville, SC today. It's near a clubhouse in a community so the maintenance supervisor requested an ID. Looking at pictures online for comparison, I think it's an anerythristic corn snake. Do you concur?


Nathan H.
South Carolina

Hi Doc,

A friend came upon this on a waterfall hike today. Can you tell what it is?

Nathan H.

What Are These Snakes?

Snake Identification Post Ground Rules

-Guesses are welcome and encouraged. Don't worry if you're not an expert, wrong guesses allow us to talk about how to distinguish between the various species and that's why I run these posts.

-If you can't explain why you think a snake is a particular species, go ahead and just say what you think it is. But otherwise please do let us all know how you identified the animal. If you're wrong, we can explain why. If you're right, this helps everyone learn how to identify snakes, which is the point of these posts.

-This is not a pop quiz, any kind of research is encouraged and I hope you will engage with other commenters to try to figure these snakes out. I will eventually chime in with my thoughts.

-Assume I know what kind of snake is in the picture. I run these posts because they are outreach opportunities. Please don't send me private e-mails with your guesses, include them below.

-Remember, the person that sent me the picture is probably reading your comments. Although it is frustrating to know that many of these snakes have been killed, these people do want to learn more about them. More snake knowledge will lead to fewer snakes being killed. Don't hate, educate.

Enjoy what you read and learn here? 

Friday, April 27, 2018

Snake Identification Challenge of the Week: The Usual Suspects

Hi Dr. Steen,

I’m a fan of your blog.  I recently purchased some property in Augusta, MO (originally from Washington DC), so I’m on a major wildlife identification kick.  In a short month, my trail cam has picked up red fox, grey fox, coyote, raccoons, possums, deer, etc.  Recently I was doing some weed eating in an area that hasn’t been touched in years and found the juvenile snake in the photo.  I carefully scooped him up and dropped him off in a pile of leaves a little deeper in the woods.  For the life of me, I can’t figure out what he is.

Thanks for any help,


 This fellow fell out of my truck door as I opened it.  I caught him and hoped he would be a rat snake based on the round pupils, but the head is too similar to his 7ft long cousin I already had a run in with.  Can you identify?  

Barbara K.

Can you tell what kind of snake this is? 

Melissa P.
North Carolina

What Are These Snakes?

Snake Identification Post Ground Rules

-Guesses are welcome and encouraged. Don't worry if you're not an expert, wrong guesses allow us to talk about how to distinguish between the various species and that's why I run these posts.

-If you can't explain why you think a snake is a particular species, go ahead and just say what you think it is. But otherwise please do let us all know how you identified the animal. If you're wrong, we can explain why. If you're right, this helps everyone learn how to identify snakes, which is the point of these posts.

-This is not a pop quiz, any kind of research is encouraged and I hope you will engage with other commenters to try to figure these snakes out. I will eventually chime in with my thoughts.

-Assume I know what kind of snake is in the picture. I run these posts because they are outreach opportunities. Please don't send me private e-mails with your guesses, include them below.

-Remember, the person that sent me the picture is probably reading your comments. Although it is frustrating to know that many of these snakes have been killed, these people do want to learn more about them. More snake knowledge will lead to fewer snakes being killed. Don't hate, educate.

Enjoy what you read and learn here? 

Tuesday, April 24, 2018

Two Unique Alabama Treasures On the Brink --Guest Post--

Flattened Musk Turtle
Take a minute and think of something unique to Alabama, something you can’t find anywhere else. What is it? Perhaps it’s tailgating on a cool Saturday afternoon after Thanksgiving, for the greatest football rivalry in the South. If you attended elementary school in Alabama, perhaps you thought of the only monument in the world to a pest insect, the Boll Weevil. Some would think of their favorite hunting or fishing honey-hole. Whatever you thought of, I doubt it is what first comes to my mind: the Black Warrior Waterdog and the Flattened Musk Turtle. In fact, most Alabamians don’t even know they exist. Native to the Upper Black Warrior River System, the Black Warrior Waterdog, Necturus alabamensis, and the Flattened Musk Turtle, Sternotherus depressus, are endemic to Alabama, meaning they only occur in Alabama and nowhere else. This alone makes these two species incredibly special, but the curious nature of the Flattened Musk Turtle and mysticism of the Black Warrior Waterdog along with the astounding beauty of the habitat in which they occur establish them as natural treasures. 

Black Warrior Waterdog (Courtesy Mark Bailey)
While conducting studies on these species over the past five years with the Alabama Natural Heritage Program at Auburn University, first as an undergraduate and now a graduate student, I have developed a deep connection and appreciation toward these species and the unique river system in which they occur. Being part of what may be the final generation of researchers to have the privilege of working with them, it is an appreciation I hope to share with you.
      The morning fog floating above the water’s surface begins to dissipate as I reach the sandy bank of the Sipsey Fork after a half-mile of kayak dragging. The steep path between bluffs was once an ATV trail. Before that, it may have been a horse and wagon trail used by settlers, and, still earlier, almost certainly was a Chickasaw Indian footpath to access the stream. Now, used only by me and the occasional deer, it is overgrown with briars and muscadine. A sliver of regret develops in the back of my mind over the decision to bring the kayak as I dread the idea of dragging 70 pounds of kayak and gear back up the 200 foot elevation gain to my car. Still though, the Sipsey Fork is uncomfortably cold to be wading in March and the feeling is quickly replaced with contentment as I begin paddling downstream along a colossal sandstone bluff overhanging the water. I steer the kayak between the steady drips of water falling 70 feet from the top of the bluff, avoiding the unpleasant sensation of cold water droplets exploding on my neck.

I paddle to my first trap which is set next to a car-sized submerged boulder. I pull the trap to find nothing but a Sloped Crayfish (Cambarus obstipus) and a couple young sunfish which I release. In the next trap, by a submerged log, I notice a dark, slender shape. My heart begins to race as I pull the trap from the water and the critter becomes visible. To my disappointment, the shape morphs into a small catfish, not my target, which is the rare aquatic salamander known as the Black Warrior Waterdog. I continue to check the rest of the traps, but they don’t yield much else. As elusive as they are, I knew the odds were against me capturing one of those foot-long salamanders.

Bizarre and majestic in their own right, Black Warrior Waterdogs, with their external gills, resemble mythical dragons of Asian lore and are quickly becoming as hard to find. They have declined drastically across their range over the past few decades due to increased sedimentation and poor water quality. Recent surveys indicate that they may have already disappeared from 80 to 95 percent of their historic range. To make matters worse, very little is known about their life history and historic abundances as few studies have been conducted on the species. Because of the lack of research, conservation of the species has been ignored. It is possible that we will lose the Black Warrior Waterdog (one of Alabama’s only two endemic amphibians) within the next couple decades.

My hope was to capture a waterdog at this site, where they hadn’t been previously sampled, to determine the extent of their range within Bankhead National Forest, but that would have to wait another year as the active season for waterdogs was drawing to a close. With traps pulled, I grab my radio telemetry receiver and antenna and turn my attention to another imperiled resident of the Sipsey Fork.

 I switch radio frequencies to a familiar wavelength. “Ping… ping…ping,” the receiver begins its metronomic tune. The volume increases as I paddle closer to the location of the transmitter. I triangulate the location to a submerged slab rock which had broken off the face of the bluff many years ago, perhaps at a time when the Chickasaw were using my path down to the stream. I scan the stream bottom while trying to hold the kayak steady in the current. Suddenly, I feel that rush of adrenaline that hunters can relate to. It’s the feeling you get in the split-second between the pattern recognition of your quarry, say a deer, in your peripheral vision and when you focus on and confirm that it is, in fact, a deer. My eyes adjust and focus in on the familiar sight of a Flattened Musk Turtle poking her yellow head with black reticulations out from under a rock.

The Flattened Musk Turtle gets its name from the uniquely flattened shape of its shell, a direct product of Alabama’s geology. Below the Fall Line, the dividing line between the Piedmont and the Coastal Plain where the Western Interior Seaway existed 100 million years ago, the Black Warrior River is mostly sandy with some small rocks and gravel. Above the Fall Line, however, the Black Warrior River Basin is a heterogeneous mix of bedrock, boulders, large slab rocks, gravel, and sand. This habitat, abundant with rock-crevices and isolated from other river systems, is where the Flattened Musk Turtle evolved.

Over hundreds of millennia, this new species of musk turtle evolved in the Upper Black Warrior that was unlike those found in the Coastal Plain or adjacent watersheds in the Piedmont. It developed a more compressed shell with a flattened carapace, reducing drag and allowing for individuals to lodge themselves deeper into crevices to escape predators and high velocity flow events - an important adaption for a river system with steep canyons that channel water creating extreme currents and exceptionally high flood levels. There were, of course, biological costs to compressing organs and living in these cold upland streams. Females took seven years to reach sexual maturity instead of five and laid half as many clutches per year. However, the wealth of food in the form of snails and great abundance of rock-crevice refuges were worth the costs.

This specialization was a good strategy as long as the streams had plenty of crevice cover. Unfortunately, that habitat has been increasingly altered over the past 70 years. Largely from extensive mountain-top removal, also known as “strip mining,” coal extraction in addition to the state government’s allowance of poor riparian zone management around some land development projects and forestry operations, sediment has inundated much of the Black Warrior Basin, filling in the crevice habitat that is so critical to the survival of the Flattened Musk Turtle. The coal mines have the double whammy of also creating long-term pollution issues that kill off aquatic mollusks (snails and mussels) that the turtles rely on for prey. Historic survey data is robust for the Flattened Musk Turtle and recent surveys indicate that they have been lost from around 70 to 90 percent of their historic range. The story of the Black Warrior Waterdog parallels that of the Flattened Musk Turtle, from their speciation in the Upper Black Warrior to their flattened body shape, and from the population declines to the issues causing them. Likewise, if we push the Black Warrior Waterdog off the cliff of extinction, the Flattened Musk Turtle, our only endemic reptile, will not be far behind. However, if we develop strategic conservation plans soon, we may save both for the price of saving one and, in the process, protect a number of other imperiled aquatic species in the Black Warrior.

This was the first time I had seen the adult female since she became inactive in November under a boulder about 40 feet upstream. I grab my GPS and begin recording data about habitat and location while she continues to watch my every move with curiosity from the comfort of her sandstone retreat. After recording the necessary data, I take a moment to admire this comical-looking turtle. Although only the size of my hand, she has been roaming this stretch of river longer than the 21 years I have been alive. I continue on, tracking other turtles and recording data before calling it a day and paddling back to the take-out. Knocking from a Pileated Woodpecker echoes across the forest as the afternoon sun dips below the bluff-line. I begin the long drag up and out of the canyon, but I can’t get my mind off that old turtle staring at me from under her rock. There is something humbling about looking into the eyes of such a creature, knowing that she has seen more sunrises than you.

This article was originally published in the magazine of the Alabama Wildlife Federation, “Alabama Wildlife,” in January 2018. The Alabama Wildlife Federation (AWF) has kindly allowed the republishing of this article, and I encourage all who care about and enjoy the incredible wildlife of the Southeast’s most biodiverse state to join AWF and support their mission to protect and educate the public aboutAlabama’s wildlife
About the Author

Joseph Jenkins, born and raised in Alabama and has had a life-long passion for amphibians and reptiles and the outdoors. He is currently working on his M.S. in Biological Sciences at Auburn University and is advised by David Steen. His thesis explores aspects of the home range, movement patterns, and habitat selection of the flattened musk turtle.